The JAWA engine is a very typical MC engine.....with high compression, and laid down on the chassis.
It's points of departure from the usual big 4 strokes is that it is a full 500 cc's.....and has 15:1 compression.
An engine, with 10% more displacement and a couple more points of compression, is going to be faster than a 450 cc engine...
other qualities of the racing program being somewhat equal.
It's not rocket science....or magic.....just more compression, more displacement than the 450's.
Most 450's have a 500cc optional jug, and piston/crankshaft.